Reaction
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Reactions can be performed in response to specific trigger. By default you can perform a single reaction in a round, though various feats increase this amount. It is usually the player's choice whether to perform a reaction, or save it for later.
All characters gain the Ready Action reaction by default. Other reactions are gained through feats and class abilities.
Threatened Area
Some reactions require creatures to be within your threatened area. You threaten all squares into which you can make an unarmed melee attack, even when it is not your turn. Generally, that means everything in all squares adjacent to your space (including diagonally).
Reactions that consist of an attack has a threatened area that matches any square that you can hit with the weapon used. One common example of this are Reach weapons, which threaten squares 10 ft. away, but not squares adjacent to you.